Tylopilus

Tylopilus Fotografia de Tylopilus felleus (boleto amargo ou boleto de vesícula) Classificação de acordo com MycoBank

Reinado	Fungi
Divisão	Basidiomycota
Subdivisão	Agaricomicotina
Aula	Agaricomicetes
Subclasse	Agaricomycetidae
Pedido	Boletales
Família	Boletaceae

Gentil

Tylopilus
P. Karst. , 1881

Tylopilus é um gênero de fungos basidiomicetos da família das Boletaceae . Existe apenas uma espécie na Europa, Tylopilus felleus . Algumas espécies foram transferidas para o gênero Rubinoboletus e o gênero Porphyrellus .

Taxonomia

Tylopilus P. Karsten On Às vezes inclui Porphyrellus , algumas espécies são transferidas para Rubinoboletus .

Descrição do esporóforo

Cutícula seca, glabra a subtomentosa. Polpa branca, imutável ou marrom clara, vermelha depois preta, ou mais raramente azul, com um sabor doce ou amargo. Hymenium adnexate, carne, em seguida, rosa branco, castanho púrpura para castanho oxidado com coloração castanha. Estipe seca, pruinosa a glabra, até reticulada, com finas crostas . Carne rosada. Cor de esporo a marrom arroxeado a marrom enferrujado. Esporos lisos, fusiformes.

Habitat

Zona temperada do Norte, América tropical montanhosa, norte da América do Sul, Leste Asiático, Sul da Ásia, Austrália, Nova Zelândia, África.

Micorrizas com Pinaceae , Fagaceae , Betulaceae , Myrtaceae , Casuarinaceae , leguminosas Cisalpinas .

Lista de espécies

De acordo com o Catálogo da Vida (28 de outubro de 2013)  :

• Tylopilus acutesquamosus
• Tylopilus alboater
• Tylopilus albofarinaceus
• Tylopilus alkalixanthus
• Tylopilus alutaceoumbrinus
• Tylopilus ammiratii
• Tylopilus appalachiensis
• Tylopilus arenarius
• Tylopilus argillaceus
• Tylopilus atratus
• Tylopilus atrobrunneus
• Tylopilus atronicotianus
• Tylopilus austrofelleus
• Tylopilus badiceps
• Tylopilus balloui
• Tylopilus beelii
• Tylopilus brachypus
• Tylopilus brevisporus
• Tylopilus brunneirubens
• Tylopilus brunneus
• Tylopilus bulbosus
• Castanoides de Tylopilus
• Tylopilus celulosus
• Tylopilus chromoreticulatus
• Tylopilus corneri
• Tylopilus costaricensis
• Tylopilus cyaneocinctus
• Tylopilus cyanescens
• Tylopilus cyanogranulifer
• Tylopilus apertado
• Tylopilus felleus
• Tylopilus ferrugineus
• Tylopilus formosus
• Tylopilus fuligineoviolaceus
• Tylopilus fumosipes
• Tylopilus funerarius
• Tylopilus gastromycetoides
• Tylopilus gomezii
• Tylopilus griseocarneus
• Tylopilus guanacastensis
• Tylopilus hondurensis
• Tylopilus hongoi
• Tylopilus humilis
• Tylopilus indecisus
• Tylopilus intermedius
• Tylopilus isabellescens
• Tylopilus jalapensis
• Tylopilus javanicus
• Tylopilus leucomycelinus
• Tylopilus lividobrunneus
• Tylopilus louisii
• Tylopilus microsporus
• Tylopilus minor
• Tylopilus mitissimus
• Tylopilus montanus
• Tylopilus montoyae
• Tylopilus nanus
• Tylopilus nebulosus
• Tylopilus neofelleus
• Tylopilus nicaraguensis
• Tylopilus niger
• Tylopilus nigricans
• Tylopilus nigropurpureus
• Tylopilus obscureviolaceus
• Tylopilus obscurus
• Tylopilus ochraceosquamosus
• Tylopilus oradivensis
• Tylopilus orsonianus
• Tylopilus otsuensis
• Tylopilus pachycephalus
• Tylopilus pakaraimensis
• Tylopilus peralbidus
• Tylopilus pernanus
• Tylopilus perplexus
• Tylopilus piniphilus
• Tylopilus pisciodorus
• Tylopilus plumbeoviolaceoides
• Tylopilus plumbeoviolaceus
• Tylopilus porphyrosporus
• Tylopilus potamogeton
• Tylopilus praeanisatus
• Pseudoscaber de Tylopilus
• Tylopilus punctatofumosus
• Tylopilus rhoadsiae
• Tylopilus rhodocônio
• Tylopilus rigens
• Tylopilus rubrobrunneus
• Tylopilus rufonigricans
• Tylopilus rugulosoreticulatus
• Tylopilus sanctae-rosae
• Tylopilus sinicus
• Tylopilus snellii
• Tylopilus striatulus
• Tylopilus suavissimus
• Tylopilus subcellulosus
• Tylopilus subfusipes
• Tylopilus subniger
• Tylopilus subunicolor
• Tylopilus subvinaceipallidus
• Tylopilus tabacinus
• Tylopilus temucensis
• Tylopilus tenuis
• Tylopilus valens
• Tylopilus variobrunneus
• Tylopilus veluticeps
• Tylopilus vinaceipallidus
• Tylopilus vinaceogriseus
• Tylopilus vinosobrunneus
• Tylopilus violaceus
• Tylopilus violatinctus
• Tylopilus virens
• Tylopilus viscidichromapes
• Tylopilus williamsii
• Tylopilus zambianus

De acordo com o Index Fungorum (28 de outubro de 2013)  :

• Tylopilus acutesquamosus Singer 1983
• Tylopilus alboater (Schwein.) Murrill 1909
• Tylopilus albofarinaceus (WF Chiu) FL Tai 1979
• Tylopilus alkalixanthus Halling & Amtoft 2003
• Tylopilus alutaceoumbrinus Hongo 1971
• Tylopilus ammiratii Thiers 1975
• Tylopilus appalachiensis Singer 1948
• Tylopilus arenarius Singer 1978
• Tylopilus areolatus (Berk.) Henn. 1898
• Tylopilus areolatus Hongo 1962
• Tylopilus argillaceus Hongo 1985
• Tylopilus atratus Ambos 1998
• Tylopilus atrobrunneus (Lj.N. Vassiljeva) Wolfe 1980
• Tylopilus atronicotianus Ambos 1998
• Tylopilus austrofelleus (Cleland) Watling & NM Greg. 1988
• Tylopilus badiceps (Peck) AH Sm. & Thiers 1971
• Tylopilus balloui (Peck) Singer 1947
• Tylopilus beelii Heinem. & Gooss.-Font. 1951
• Tylopilus brachypus Singer, J. García & LD Gómez 1991
• Tylopilus brevisporus Watling e TH Li 1999
• Tylopilus brunneirubens (Corner) Watling & E. Turnbull 1994
• Tylopilus brunneus (McNabb) Wolfe 1980
• Tylopilus bulbosus Halling & GM Muell. 2001
• Tylopilus castanoides Har. Takah. 2002
• Tylopilus cellulosus Singer 1947
• Tylopilus chromoreticulatus Wolfe & Bougher 1993
• Tylopilus conspicuocystidiata Watling & E. Turnbull 1994
• Tylopilus corneri Singer, J. García & LD Gómez 1991
• Tylopilus costaricensis Singer e LD Gómez 1991
• Tylopilus cyaneocinctus (Singer) Grund & KA Harrison 1976
• Tylopilus cyanescens TH Li & Watling 1999
• Tylopilus cyanogranulifer TH Li, Watling & NM Greg. 1999
• Tylopilus exiguus TW Henkel 1999
• Tylopilus felleus (Bull.) P. Karst. 1881
• Tylopilus ferrugineus (Frost) Singer 1947
• Tylopilus formosus G. Stev. 1962
• Tylopilus fuligineoviolaceus Har. Takah. 2007
• Tylopilus fumosipes (Peck) AH Sm. & Thiers 1971
• Tylopilus funerarius (Massee) Pegler & TWK Young 1981
• Tylopilus fuscobrunneus Watling 1989
• Tylopilus gastromycetoides TH Li 1999
• Tylopilus gomezii Singer 1991
• Tylopilus griseocarneus Wolfe & Halling 1989
• Tylopilus guanacastensis Singer 1983
• Tylopilus hondurensis Singer e MH Ivory 1983
• Tylopilus hongoi Wolfe & Bougher 1993
• Tylopilus humilis Thiers 1967
• Tylopilus indecisus (Peck) Murrill 1909
• Tylopilus intermedius AH Sm. & Thiers 1971
• Tylopilus isabellescens Murrill 1946
• Tylopilus jalapensis Singer e J. García 1991
• Tylopilus javanicus Henn. 1899
• Tylopilus leucomycelinus (Singer & MH Ivory) R. Flores & Simonini 2000
• Tylopilus lividobrunneus Singer 1973
• Tylopilus louisii Heinem. 1964
• Tylopilus microsporus SZ Fu, QB Wang e YJ Yao 2006
• Tylopilus minor Singer 1945
• Tylopilus mitissimus Singer e LD Gómez 1991
• Tylopilus montanus Singer 1989
• Tylopilus montoyae Singer e J. García 1991
• Tylopilus nanus (Massee) Singer 1975
• Tylopilus nebulosus (Peck) Wolfe 1980
• Tylopilus neofelleus Hongo 1967
• Tylopilus nicaraguensis Singer e MH Ivory 1983
• Tylopilus niger (Heinem. & Gooss.-Font.) Wolfe 1980
• Tylopilus nigricans Singer 1947
• Tylopilus nigropurpureus (Canto) Hongo 1973
• Tylopilus obscureviolaceus Har. Takah. 2004
• Tylopilus obscurus Halling 1989
• Tylopilus ochraceosquamosus Watling 1994
• Tylopilus olivaceoporus Watling 1989
• Tylopilus oradivensis Osmundson e Halling 2010
• Tylopilus orsonianus Fulgenzi & TW Henkel 2007
• Tylopilus otsuensis Hongo 1966
• Tylopilus pachycephalus (Massee) Singer 1983
• Tylopilus pakaraimensis TW Henkel 2001
• Tylopilus peralbidus (Snell & Beardslee) Murrill 1938
• Tylopilus pernanus (Pat. & CF Baker) Watling 2000
• Tylopilus perplexus Watling 1994
• Tylopilus piniphilus Wolfe & Bougher 1993
• Tylopilus pisciodorus (Murrill) Murrill 1948
• Tylopilus plumbeoviolaceoides TH Li, B. Song e YH Shen 2002
• Tylopilus plumbeoviolaceus (Snell & EA Dick) Snell & EA Dick 1941
• Tylopilus porphyrosporus (Fr. & Hök) AH Sm. & Thiers 1971
• Tylopilus potamogeton Singer 1978
• Tylopilus praeanisatus (Murrill) Murrill 1948
• Tylopilus pseudoscaber Secr. ex AH Sm. & Thiers 1968
• Tylopilus pumilus (Saut.) Henn. 1898
• Tylopilus punctatofumosus (WF Chiu) FL Tai 1979
• Tylopilus rhoadsiae (Murrill) Murrill 1944
• Tylopilus rhodoconius (Singer) TJ Baroni, Both & Bessette 1998
• Tylopilus rigens Hongo 1979
• Tylopilus rubrobrunneus Mazzer & AH Sm. 1967
• Tylopilus rufonigricans TW Henkel 1999
• Tylopilus rugulosoreticulatus Hongo 1979
• Tylopilus sanctae-rosae Singer 1983
• Tylopilus shaukatii Gardezi 2007
• Tylopilus sinicus (WF Chiu) FL Tai 1979
• Tylopilus snellii Wolfe 1983
• Tylopilus sordidus (Frost) AH Sm. & Thiers 1968
• Tylopilus striatulus Heinem. 1951
• Tylopilus suavissimus Heinem. & Gooss.-Font. 1951
• Tylopilus subcellulosus Singer e J. García 1991
• Tylopilus subfusipes AH Sm. 1973
• Tylopilus subniger Singer, J. García & LD Gómez 1991
• Tylopilus subunicolor EA Dick & Snell 1961
• Tylopilus subvinaceipallidus TH Li & Watling 1999
• Tylopilus tabacinus (Peck) Singer 1944
• Tylopilus temucensis Palfner 2005
• Tylopilus tenuis Heinem. 1951
• Tylopilus valens (Canto) Hongo e Nagas. 1976
• Tylopilus variobrunneus Roody, AR Bessette & Bessette 1998
• Tylopilus velatus (Rostr.) FL Tai 1979
• Tylopilus veluticeps (Pat. & CF Baker) Singer 1947
• Tylopilus vinaceipallidus (Corner) TW Henkel 1999
• Tylopilus vinaceogriseus Singer, J. García & LD Gómez 1991
• Tylopilus vinosobrunneus Hongo 1979
• Tylopilus violaceus Heinem. 1951
• Tylopilus violatinctus TJ Baroni & Both 1998
• Tylopilus virens (WF Chiu) Hongo 1964
• Tylopilus viscidichromapes Watling & TH Li 1999
• Tylopilus williamsii Singer e J. García 1991
• Tylopilus zambianus Watling 1994

De acordo com o NCBI (28 de outubro de 2013)  :

• Tylopilus alboater
• Tylopilus appalachiensis
• Tylopilus atronicotianus
• Tylopilus badiceps
• Tylopilus balloui
• Tylopilus cartagoensis
• Tylopilus castaneiceps
• Tylopilus apertado
• Tylopilus felleus
• Tylopilus ferrugineus
• Tylopilus formosus
• Tylopilus indecisus
• Tylopilus intermedius
• Tylopilus leucomycelinus
• Tylopilus neofelleus
• Tylopilus palumanus
• Tylopilus pernanus
• Tylopilus plumbeoviolaceoides
• Tylopilus plumbeoviolaceus
• Tylopilus porphyrosporus
• Tylopilus propriorichromapes
• Pseudoscaber de Tylopilus
• Tylopilus queenslandianus
• Tylopilus rhoadsiae
• Tylopilus rubrobrunneus
• Tylopilus rufonigricans
• Tylopilus tabacinus
• Tylopilus variobrunneus
• Tylopilus vinosobrunneus
• Tylopilus violatinctus
• Tylopilus virens

Notas e referências

1. Roskov Y., Ower G., Orrell T., Nicolson D., Bailly N., Kirk PM, Bourgoin T., DeWalt RE, Decock W., van Nieukerken EJ, Penev L. (eds.) (2020). Species 2000 & ITIS Catalog of Life , 2020-12-01. Recurso digital em www.catalogueoflife.org . Species 2000: Naturalis, Leiden, Holanda. ISSN 2405-8858, acessado em 28 de outubro de 2013
2. Index Fungorum , acessado em 28 de outubro de 2013
3. NCBI , acessado em 28 de outubro de 2013

links externos

• (in) Catálogo de referência da vida  : Tylopilus (acessado12 de dezembro de 2020)
• (pt) Referência do índice Fungorum  : Tylopilus P. Karst. 1881 ( + lista de espécies ) ( + MycoBank ) (consultado em28 de outubro de 2013)
• (en) Referência do MycoBank  : Tylopilus P. Karst. (consultou o28 de outubro de 2013)
• (pt) Referência do NCBI  : Tylopilus ( táxons incluídos ) (consultado em28 de outubro de 2013)
• (en) uBio referência  : Tylopilus P. Karst. 1881 (consultado em28 de outubro de 2013)